In Volume 18, Ladon joins in the attack on the Heaven where he teams up with three Grendel clones but was ultimately defeated by the joint effort of Cao Cao and the Gremory Group, and was later sealed into a detached Boosted Gear jewel. He fought and successfully trapped Issei using multiple barriers until he was called back by Euclid using a Dragon Gate. Ladon later appeared at the Southwest region of Auros together with Grendel. Rizevim told D×D that the barrier surrounding Auros and Agreas was created by Ladon, assisted by Euclid's replica Boosted Gear.
Ladon first appeared in Volume 17 alongside Rizevim Livan Lucifer, Euclid Lucifuge, and Azi Dahaka with Rizevim explaining his plans.
Plot The Legend of Oppai Dragon and his Lively Companions He was later revived by the Khaos Brigade sometime before Volume 17. In the past, Ladon was in charge of guarding the Golden Apples of Hesperides and was killed by the original Heracles to steal the golden apples as part of his labors. Ladon appears to be prideful in the strength of his barriers, wanting to "test" them against strong opponents such as Rias' Power of Destruction, or Issei's Cardinal Crimson Promotion. Ladon is a tree-like Dragon with red eyes.
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